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Glutathione S -Transferases (GSTs) GSH has many biological roles, including (1) defense against reactive oxygen species and free radicals, (2) defense against reactive electrophiles, though the action of GSTs (3) carrier of cysteine in a less reactive form, (4) cofactor in several enzyme reactions, (5) participation in eicosanoid biology, (6) maintenance of cellular redox state, and (7) transport of cysteine (e.g., Taniguchi et al., 1989)